400年前,沃克家唯一的一个孩子从海盗“辛格兄弟会”的屠刀之下死里逃生,并意外获得了一件刀枪不入的战衣,从此,专门惩奸除恶的“幻影侠”诞生了。一晃眼已是二十一世纪,克里斯(莱安·卡勒斯 Ryan Carnes 饰)是沃克家的第二十一代传人,为了躲避辛格兄弟会的追杀,他不得不离开了家乡,跟随养父母来到了美国。 长大之后,克里斯成为了一名法学院的学生,然而,将做律师视为毕生理想的他怎么也无法想到的是,自己即将肩负起整个家族的使命,成为新一代“幻影侠”。在颅骨洞里,克里斯得到了家族中代代相传的战衣,与此同时,辛格兄弟会的成员们杀害了克里斯的养父母,还将下一个目标放在了克里斯的女友蕾妮(卡梅隆·古德曼 Cameron Goodman 饰)的身上。
At the beginning of 2259, Captain John Sheridan replaces Sinclair as the military governor of the station. He and the command staff learn that the death of President Santiago was actually an assassination masterminded by Vice President Clark (who assumed the Presidency upon Santiago's death). A conflict develops between the Babylon 5 command staff and the Psi Corps, an increasingly autocratic organization which oversees and controls the lives of human telepaths. Commander Ivanova, the second-in-command of the station, is secretly a latent telepath who has illicitly avoided registering with the Psi Corps. The Shadows, an ancient and extremely powerful race who have recently emerged from hibernation, are revealed to be the cause of a variety of mysterious and disturbing events, including the attack on the Narn outpost at the end of 2258. Centauri Ambassador Londo Mollari unknowingly enlists their aid through his association with the mysterious Mr. Morden in the ongoing conflict with the Narn. The elderly and ailing Centauri emperor, long an advocate of reconciliation with the Narn, dies suddenly while visiting Babylon 5. A number of conspirators, including Londo Mollari and Refa, take control of the Centauri government by assassinating their opponents and placing the late emperor's unstable nephew on the throne. Their first act is to start open aggression against the Narn. After a full-scale war breaks out, the Centauri eventually conquer Narn in a brutal attack involving mass drivers, outlawed weapons of mass destruction. Towards the end of the year, the Clark administration begins to show increasingly totalitarian characteristics, clamping down on dissent and restricting freedom of speech. The Vorlons are revealed to be the basis of legends about angels on various worlds, including Earth, and are the ancient enemies of the Shadows. They enlist the aid of Sheridan and the Babylon 5 command staff in the struggle against the Shadows.